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Funding: The primary funder, Vertex Pharmaceuticals, Inc., provided a grant to investigate MRI biomarkers of myelination which supported experimental costs and partial salary support for authors AOC, JP, IT, and WR, but did not have any additional role in the study design, data collection and analysis, decision to publish, or preparation of the manuscript

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The arterial endothelial cells uniquely expressed Alpl and Hey1 , the capillary endothelial cells strongly expressed Lpl , and the venous endothelial cells expressed Vwf , Hif1a , Icam1 , Lrg1 and Aplnr (Extended Data Fig