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In colon cancer (HCT 116) cells, emodin reduces VEGF secretion and VEGFR1/2 protein expression by down-regulating Acyl-CoA synthetase long chain family member 4 (ACSL4) [468], while in liver cancer, emodin inhibits tumorigenesis through concurrently suppressing the VEGFR2-AKT-ERK1/2 signaling pathway, and inhibiting the protein expression of SMAD2/4 by enhancing miR-34a level in Hep G2 cells [469]

In the nervous system, the pathway interacts with NMDA receptor activity and neuropeptide release, thereby modulating pain signaling at both peripheral and central sites